I was on the fence between a GX460 and a 4runner and bought the 4runner for a number of reasons - primarily that our Lexus dealer in Utah is HORRIBLE - so bad that I swore I'd never own another Lexus, but then my wife ended up with another one - RX350 we bought in another state- with a caveat that it gets serviced at Toyota, not the Lexus dealer.
That said, I have a '15 4runner Trail with crawl control, so I can comment to some degree. If you're serious about offroading none of these are what you want - they're all tanks. The 4runner is not light by any means, but still weighs in around 800lbs lighter than the GX = better offroad, but no heated steering wheel. Also the 4runner trail seats fold down about 10" lower than the GX460 or 4runner limited (which has basically the same interior as the GX with rear seats that don't fold all the way down). And being 800ish lbs lighter, the highway performance is a lot closer than you might think with the v6 - about 0.5 seconds to 60 slower. Road noise is comparable - both are very quiet.
As for lockers vs crawl control - I'll take mechanical lockers any day. No matter how good traction control is, it's not as good at going the last mile offroad as real lockers, but it might be better at cruising around logging roads. The reason is that crawl control still requires wheel slip to engage, and that means you've lost static friction and now have kinetic friction on the tire that slips. Static friction is much greater in most situations than kinetic friction. So with crawl control you can achieve essentially the maximum static friction on the tire with most traction and kinetic friction with the other. With a locker you can get the absolute maximum static friction from both tires on the axle. So for me, ARBs would be my choice over crawl control - or any other traction control system.
As far as the cruise control functionality, low range in 1st gear does a pretty good job. It's not as good as a manual transmission for maintaining steady speed over rough obstacles, but does a good enough job that I don't think you need crawl control. You might find that the atrac system and low range is good enough without lockers for anywhere you're realistically going to try to go in a 5,500lb luxury suv.
Finally - I'd start by adding a winch if it were me. Then find out if it goes where you want. If not, start building to be what you want. No crawl control wouldn't be a deal breaker for me.