My "O-ring" seals were definitely also profiled (not a traditional round O-ring). I ordered them from Mid Atlantic Toyota Parts, an online Toyota parts house, so I can confirm that they are stock Toyota O-rings. When I did a parts search, they appeared to be universal to all AISIN locking hubs (FJ40, FJ60, Mini Truck, Suzuki Samurai, etc.) I did not pay any attention to the orientation when I installed them, I just threw them on and buttoned the dial back up. Maybe I got lucky? I hesitate to think that's the case given that my truck stubbornly fights me every time I try to perform even the most basic maintenance, but maybe everyone gets one streak of luck...
I'm totally at a loss here. My original rings were stretched and dry rotted, but I reassembled the dials with them at one point to test fit things, and they still fit and worked. Final reassembly was done with the new rings, and everything still fit, and worked properly, lubed or dry. The C-clip was a little bit of a pain to get seated, but only so much as every other C-clip is a pain to get seated...I think I went through both dials, cleaned, lubed, new seals, and clips reinstalled, in a half hour...it shouldn't be this hard!
I can't imagine there is a similarly conflicting burr on both dials that would prevent proper seating - that would be far too coincidental. It seems that you are 100% certain that those are the same C-clips that were installed there previously (and I believe you are correct). Looking at mine, it's the only C-clip with the cut-outs for the pliers to grab...This seems to point to the O-ring/gasket, but if this was the case, I would imagine they would still reassemble with the old gaskets.
The only thing I can think to try is clearancing the rubber seal with some sand paper to see if you can minimally reduce the outer diameter in order to make it fit...Of course, then you might not get a good seal, either...
I really don't know...I'm frustrated FOR you!
For what it's worth, here are some shots of mine, during disassembly - unfortunately it doesn't look like I have a shot of the O-ring seated, but in case it provides any insight:
And a link to my thread where I reassembled mine:
https://forum.ih8mud.com/threads/th...in-refurbishing-a-1975-chevota.725833/page-18
I will try to remember to search my photo archive when I get home tonight to see if I have any additional hub shots that I didn't upload that might provide some more info...