Base models vs Base models, sure, but once you start adding in different options and the upper level trims, the differences aren't as cut and dried.
The Land Cruiser (PRADO) First Edition is $500 less than the GX550 Overtrail+ (we don't have pricing on the 4Runner yet, but I am betting that the 4Runner TRD Pro or Trailhunter will be similarly priced).
The GX550 isn't hybrid (and thats actually a feature I want, personally) but I am seeing many people bemoaning the fact that the Land Cruiser (PRADO) is a hybrid. A GX550 hybrid would be my choice, but I suspect that truck will be getting to LX600 money.
I am also not sure that the full time 4wd option is much oif a selling point when part time conversion kits like this actually exist:
Part Time Conversion Kit
cruiserteq.com
I just don't see the Land Cruiser (PRADO) as being something that will be available in the US market for very long when I think the 4Runner and GX550 will out sell it by significant margin when comparing apples to apples feature sets. The Land Cruiser has always been the 'premium' offering from Toyota, but there is less to differentiate it from their other products to make it an appealing vehicle for many buyers, including this particular 'Cruiserhead.