@Dan Lee We have both in my garage, see:
Do people think you drive a Highlander? 2008 LC 200 and 2012 Highlander Limited AWD
To quote Inigo Montoya "Let me explain... No, there is too much. Let me sum up. ... "
The Highlander AWD is a capable off-roader "softroader". Don't expect too much articulation or flex compared to the KDSS system of the LC. I have taken ours on the ranch and state parks with no issues. Think along the lines of light off road duty. Could you do more? probably. But that's not what I choose to use it for. I will have to look at the controls again: but I know it has DAC and 2nd start "Snow mode". I can check on a center locker control also. As others have said, it is built on a Camry platform. The AWD system is different than the 4WD sytem of the LC, and has no lo range. To summarize, there's a reason they don't build sliders for a Highlander.
And regarding cup holders: It has TONS of them. Where the LC has crawl control and ECT controls, the Highlander has cup holders. Where the rear pass climate controls are in the console, below that are more cup holders. Everywhere the LC has cup holders, the Highlander also has them, then adds more cup holders. To summarize, the Highlander is designed for more mass market appeal and suburban utility.
Rear lift gate is powered and remote controlled on the Highlander. Although it isn't split like the LC, it does allow for the window glass to be opened, which I like. There is also a pair of levers in the rear of the cargo area that allow you to drop the 2nd row seats into the floor. When combined with the 3rd row which also folds flat, the layout is considerable different from jump seat 3rd and a tumbled forward 2nd row. To summarize, the Highlander has some "nice to haves" but I'm not trading in the LC any time soon.
I am happy to take pics of anything you'd like regarding the differences and/or measurements between the two if it will help in your research.